Topographical constraints on competition for territories.

In homogeneous habitats, a female choosing a high quality territory should be more likely to get a high quality male also, and so it would be sufficient for her to assess only territory quality. In habitats where topography is important in determining territory boundaries females choosing on the basis of territory quality are less likely to obtain a mate of comparable quality. In these habitats, it may be optimal for a female to choose her mate on the basis of male quality as well as territory quality.-from Authors